The specificities of temperature regime of automorphic clayey soils forming under the suffruticous and shrub vegetation within the zone of tundra and forest tundra in the European North-East were studied. As the objects of investigation we chose the organic cryometamorphic soils and cryometamorphic gleezems; in the both soil types the CRM cryometamorphic horizon is developed. The soils are formed in conditions of long-termed seasonal freezing at the absence (deep occurrence) of the permafrost rocks. The dynamics near the zero temperatures (zero curtains) is characterized. The hypothesis, concerning the role of zero curtains in the sustaining of the specific angular-grainy structure within the mass of cryometamorphic horizons is formulated. The mass of cryometamorphic horizons and the depth of present-day zero curtains, which observed at the long-term seasonal soil freezing, correlate to each other. The impact of suffruticous and shrub vegetation on the specificities of winter and summer soil temperature regime is determined. We discovered that the main differences between the soils developing under suffruticous and shrub vegetation tundras are stipulated by the different intensity of the snow accumulation within these areas. The soils that are developed under the shrub vegetation are warmer than soils developed under the suffruticous tundra, where permafrost may occur at the depth of 2-3 cm. In general, seasonaly freezing tundra soils are located in the middle of the range of the automorphic clay loamy soils in the tundra-taiga ecotone of European North-East of Russia, and occupy the niche between permafrost tundra and non-permafrost north taiga soils.